1 Department of Pathology and Neurosciences, University of Dundee, Ninewells Medical School and Hospital, Dundee, DD1 9SY, UK.
2 Ontario Cancer Institute, University of Toronto, 610 University Avenue, Toronto, ON, M5G 2M9, Canada.
3 Pacific Northwest Research Institute, 720 Broadway, Seattle, WA, 98122, USA.
4 Division of Biological Chemistry, School of Life Sciences, University of Dundee, Dundee, DD1 4EH, UK.
5 Department of Pathology and Neurosciences, University of Dundee, Ninewells Medical School and Hospital, Dundee, DD1 9SY, UK. c.d.sutherland@dundee.ac.uk.