1 Laboratory of Amphibian Systematics and Evolutionary Research, College of Biology & Environment, Nanjing Forestry University, 159 Longpan Rd, Nanjing 210037, People's Republic of China.
2 Institut de Systématique, Evolution, Biodiversité, Muséum national d'Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 45 rue Buffon, CP 50, Paris 75005, France.
3 Joint Russian-Vietnamese Tropical Research and Technological Center, 64 Nguyen Van Huyen, Cau Giay, Hanoi 122000, Vietnam.
4 Department of Vertebrate Zoology, Lomonosov Moscow State University, 1/12 Leninskiye Gory, Moscow 119234, Russia.
5 Department of Zoology, Comenius University in Bratislava, Mlynská dolina, Ilkovičova 6, Bratislava 842 15, Slovakia.