1 Wellcome Sanger Institute, Cambridge, UK. mh32@sanger.ac.uk.
2 Biosciences Institute, Newcastle University, Newcastle upon Tyne, UK. mh32@sanger.ac.uk.
3 Department of Dermatology and NIHR Newcastle Biomedical Research Centre, Newcastle Hospitals NHS Foundation Trust, Newcastle upon Tyne, UK. mh32@sanger.ac.uk.
4 Wellcome Sanger Institute, Cambridge, UK.
5 Wellcome Sanger Institute, Cambridge, UK. st9@sanger.ac.uk.
6 Theory of Condensed Matter Group, Department of Physics, Cavendish Laboratory, University of Cambridge, Cambridge, UK. st9@sanger.ac.uk.