1 Department of Biomedical Sciences, Seoul National University Graduate School, Seoul, Korea (complete list of author affiliations is available in Appendix S1; see Supporting Information).
2 Department of Pathology, Seoul National University College of Medicine, Seoul, Korea.
3 Department of Dermatology, CHA Bundang Medical Center, CHA University, Seongnam, Korea.
4 Department of Dermatology, SMG-SNU Boramae Medical Center, Seoul, Korea.
5 Department of Dermatology, Seoul National University College of Medicine, Seoul, Korea.
6 Institute of Human-Environment Interface Biology, Medical Research Center, Seoul National University, Seoul, Korea.
7 Department of Pathology, CHA Bundang Medical Center, CHA University, Seongnam, Korea.
8 Laboratory for Investigative Dermatology, The Rockefeller University, New York, NY, USA.
9 Dermatology Section, Veterans Affairs Northern California Health Care System, Mather, CA, USA.
10 Department of Dermatology, University of California Davis, Sacramento, CA, USA.
11 Department of Microbiology and Immunology, Seoul National University College of Medicine, Seoul, Korea.
12 Cancer Research Institute, College of Medicine, Seoul National University, Seoul, Korea.
13 Department of Dermatology, Seoul National University Hospital, Seoul, Korea.
14 Genomic Medicine Institute, Medical Research Center, Seoul National University, Seoul, Korea.