1 Institut Pasteur, Immunoregulation Unit and Department of Immunology, Université de Paris, Paris, France.
2 Institute of Rheumatic and Musculoskeletal Medicine, University of Leeds Leeds, UK.
3 Institut Pasteur, Immunoregulation Unit and Department of Immunology, Université de Paris and Service de Rhumatologie Hôpital Cochin Port Royal, AP-HP, Paris, France.