1 CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, 650201 Kunming, Yunnan, China.
2 Kunming College of Life Science, University of Chinese Academy of Sciences, 650204 Kunming, Yunnan, China.
3 Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences, 650201 Kunming, Yunnan, China.
4 Flower Research Institute, Yunnan Agricultural Academy of Sciences, Kunming 650231, China.