1 Cell Biology Center, Institute of Innovative Research, Tokyo Institute of Technology, 4259-S2-12 Nagatsuta-cho, Midori-ku, Yokohama, 226-8503, Japan.
2 Tokyo Tech World Research Hub Initiative (WRHI), Institute of Innovative Research, Tokyo Institute of Technology, Yokohama, Japan.
3 Monash Biomedicine Discovery Institute, Department of Biochemistry and Molecular Biology, Monash University, Clayton campus, VIC, 3800, Australia.
4 Cell Biology Center, Institute of Innovative Research, Tokyo Institute of Technology, 4259-S2-12 Nagatsuta-cho, Midori-ku, Yokohama, 226-8503, Japan. yohsumi@iri.titech.ac.jp.