The structure of a potassium-selective ion channel reveals a hydrophobic gate regulating ion permeation

IUCrJ. 2020 Jul 25;7(Pt 5):835-843. doi: 10.1107/S2052252520008271. eCollection 2020 Sep 1.

Abstract

Protein dynamics are essential to function. One example of this is the various gating mechanisms within ion channels, which are transmembrane proteins that act as gateways into the cell. Typical ion channels switch between an open and closed state via a conformational transition which is often triggered by an external stimulus, such as ligand binding or pH and voltage differences. The atomic resolution structure of a potassium-selective ion channel named NaK2K has allowed us to observe that a hydro-phobic residue at the bottom of the selectivity filter, Phe92, appears in dual conformations. One of the two conformations of Phe92 restricts the diameter of the exit pore around the selectivity filter, limiting ion flow through the channel, while the other conformation of Phe92 provides a larger-diameter exit pore from the selectivity filter. Thus, it can be concluded that Phe92 acts as a hydro-phobic gate, regulating the flow of ions through the selectivity filter.

Keywords: X-ray crystallography; conformational transitions; gating; potassium ion channels.

Grants and funding

This work was funded by Advanced Research Projects Agency - Energy grants DE-AC02-06CH11357 and DE-AC02-05CH11231. Foundation for the National Institutes of Health grants GM063210 and R01-GM071939. Deutsche Forschungsgemeinschaft grant .