1 Department of Pathology, Mammalian Molecular Genetics Group, University of Cambridge, Tennis Court Road, Cambridge CB2 1QP, UK.
2 RC Centre of Excellence in Biotechnology and Development, Monash Institute for Medical Research, Monash University, Clayton, Melbourne, Victoria 3168, Australia.
3 Division of Stem Cell Research and Developmental Genetics, MRC National Institute for Medical Research, The Ridgeway, Mill Hill, London NW7 1AA, UK.