Abstract
Reprogramming somatic cells to pluripotency by Oct4, Sox2, Klf4, and Myc represent a paradigm for cell fate determination. Here, we report a combination of Jdp2, Jhdm1b, Mkk6, Glis1, Nanog, Essrb, and Sall4 (7F) that reprogram mouse embryonic fibroblasts or MEFs to chimera competent induced pluripotent stem cells (iPSCs) efficiently. RNA sequencing (RNA-seq) and ATAC-seq reveal distinct mechanisms for 7F induction of pluripotency. Dropout experiments further reveal a highly cooperative process among 7F to dynamically close and open chromatin loci that encode a network of transcription factors to mediate reprogramming. These results establish an alternative paradigm for reprogramming that may be useful for analyzing cell fate control.
Copyright © 2019 The Authors. Published by Elsevier Inc. All rights reserved.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Cell Differentiation / genetics
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Cells, Cultured
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Cellular Reprogramming / genetics*
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Chimera / genetics
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DNA-Binding Proteins / genetics
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DNA-Binding Proteins / metabolism*
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Embryonic Stem Cells / cytology
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Embryonic Stem Cells / metabolism
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Euchromatin / genetics
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Euchromatin / metabolism
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F-Box Proteins / genetics
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F-Box Proteins / metabolism*
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Fibroblasts / cytology
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Fibroblasts / metabolism
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Heterochromatin / genetics
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Heterochromatin / metabolism
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Induced Pluripotent Stem Cells / cytology*
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Induced Pluripotent Stem Cells / metabolism
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Jumonji Domain-Containing Histone Demethylases / genetics
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Jumonji Domain-Containing Histone Demethylases / metabolism*
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Kruppel-Like Factor 4
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MAP Kinase Kinase 6 / genetics
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MAP Kinase Kinase 6 / metabolism*
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Mice
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Mice, Inbred C57BL
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Mice, Transgenic
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Nanog Homeobox Protein / genetics
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Nanog Homeobox Protein / metabolism*
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RNA-Seq
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Repressor Proteins / genetics
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Repressor Proteins / metabolism*
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Transcription Factors / genetics
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Transcription Factors / metabolism*
Substances
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DNA-Binding Proteins
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Euchromatin
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F-Box Proteins
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Glis1 protein, mouse
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Heterochromatin
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Jundp2 protein, mouse
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Klf4 protein, mouse
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Kruppel-Like Factor 4
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Nanog Homeobox Protein
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Nanog protein, mouse
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Repressor Proteins
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Sall4 protein, mouse
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Transcription Factors
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Jumonji Domain-Containing Histone Demethylases
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Kdm2b protein, mouse
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MAP Kinase Kinase 6
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Map2k6 protein, mouse