1 Shandong Provincial Key Laboratory of Plant Stress, College of Life Science, Shandong Normal University, Jinan, 250014, China.
2 The University of Queensland, School of Agriculture and Food Sciences, QLD, St Lucia, 4072, Australia.
3 The State Key Laboratory of Plant Cell and Chromosome Engineering, and Center for Genome Editing, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, 100101, China. cxgao@genetics.ac.cn.
4 University of Chinese Academy of Sciences, Beijing, 100049, China. cxgao@genetics.ac.cn.