Exchange Proteins Directly Activated by cAMP and Their Roles in Respiratory Syncytial Virus Infection

J Virol. 2018 Oct 29;92(22):e01200-18. doi: 10.1128/JVI.01200-18. Print 2018 Nov 15.

Abstract

Respiratory syncytial virus (RSV) is the leading cause of respiratory infection in young children and high-risk adults. However, a specific treatment for this viral infection is not currently available. In this study, we discovered that an exchange protein directly activated by cyclic AMP (EPAC) can serve as a potential therapeutic target for RSV. In both lower and upper epithelial cells, treatment with EPAC inhibitor (ESI-09), but not protein kinase A inhibitor (H89), significantly inhibits RSV replication and proinflammatory cytokine/chemokine induction. In addition, RSV-activated transcriptional factors belonging to the NF-κB and IRF families are also suppressed by ESI-09. Through isoform-specific gene knockdown, we found that EPAC2, but not EPAC1, plays a dominant role in controlling RSV replication and virus-induced host responses. Experiments using both EPAC2 knockout and EPAC2-specific inhibitor support such roles of EPAC2. Therefore, EPAC2 is a promising therapeutic target to regulate RSV replication and associated inflammation.IMPORTANCE RSV is a serious public health problem, as it is associated with bronchiolitis, pneumonia, and asthma exacerbations. Currently no effective treatment or vaccine is available, and many molecular mechanisms regarding RSV-induced lung disease are still significantly unknown. This project aims to elucidate an important and novel function of a protein, called EPAC2, in RSV replication and innate inflammatory responses. Our results should provide an important insight into the development of new pharmacologic strategies against RSV infection, thereby reducing RSV-associated morbidity and mortality.

Keywords: EPAC; EPAC2; RSV; inflammation; replication; replication and immune response.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • A549 Cells
  • Animals
  • Cell Line
  • Chemokines / immunology
  • Cyclic AMP-Dependent Protein Kinases / antagonists & inhibitors
  • Guanine Nucleotide Exchange Factors / antagonists & inhibitors*
  • Guanine Nucleotide Exchange Factors / genetics*
  • Humans
  • Hydrazones / pharmacology
  • Isoquinolines / pharmacology
  • Isoxazoles / pharmacology
  • Mice
  • NF-kappa B / antagonists & inhibitors
  • RNA Interference
  • RNA, Small Interfering / genetics
  • Respiratory Syncytial Virus Infections / virology
  • Respiratory Syncytial Virus, Human / genetics*
  • Respiratory Syncytial Virus, Human / growth & development*
  • Sulfonamides / pharmacology
  • Virus Replication / physiology*

Substances

  • 3-(5-tert-butylisoxazol-3-yl)-2-((3-chlorophenyl)hydrazono)-3-oxopropionitrile
  • Chemokines
  • Guanine Nucleotide Exchange Factors
  • Hydrazones
  • Isoquinolines
  • Isoxazoles
  • NF-kappa B
  • RAPGEF3 protein, human
  • RAPGEF4 protein, human
  • RNA, Small Interfering
  • Sulfonamides
  • Cyclic AMP-Dependent Protein Kinases
  • N-(2-(4-bromocinnamylamino)ethyl)-5-isoquinolinesulfonamide