1 Department of Cell and Tissue Biology and Program in Craniofacial Biology, Institute for Human Genetics, University of California at San Francisco, San Francisco, CA 94143, USA. Electronic address: Jeffrey.Bush@ucsf.edu.
2 Department of Biosciences and Nutrition, Karolinska Institutet, Stockholm, Sweden; Department of Cell and Molecular biology, Karolinska Institutet, Stockholm, Sweden. Electronic address: emma.andersson@ki.se.