Pericentric H3K9me3 Formation by HP1 Interaction-defective Histone Methyltransferase Suv39h1

Cell Struct Funct. 2016 Dec 3;41(2):145-152. doi: 10.1247/csf.16013. Epub 2016 Oct 12.

Abstract

Pericentric regions form epigenetically organized, silent heterochromatin structures that accumulate histone H3 lysine 9 tri-methylation (H3K9me3) and heterochromatin protein 1 (HP1), a methylated H3K9-binding protein. At pericentric regions, Suv39h is the major enzyme that generates H3K9me3. Suv39h also interacts directly with HP1. However, the importance of HP1 interaction for Suv39h-mediated H3K9me3 formation at the pericentromere is not well characterized. To address this question, we introduced HP1 binding-defective, N-terminally truncated mouse Suv39h1 (ΔN) into Suv39h-deficient cells. Pericentric H3K9me3-positive cells were not detected by endogenous-level expression of ΔN. Notably, ΔN could induce pericentric accumulation of H3K9me3 as wild type Suv39h1 did if it was overexpressed. These findings demonstrate that the N-terminal region of Suv39h1, presumably via HP1-Suv39h1 interaction, is required for Suv39h1-mediated pericentric H3K9me3 formation, but can be overridden if Suv39h1 is overproduced, indicating that Suv39h1-mediated heterochromatin formation is controlled by multiple modules, including HP1.

MeSH terms

  • Animals
  • Blotting, Western
  • Chromobox Protein Homolog 5
  • Chromosomal Proteins, Non-Histone / metabolism*
  • HEK293 Cells
  • Heterochromatin / metabolism
  • Histones / metabolism*
  • Humans
  • Methylation
  • Methyltransferases / chemistry
  • Methyltransferases / genetics
  • Methyltransferases / metabolism*
  • Mice
  • Microscopy, Fluorescence
  • Phenotype
  • Plasmids / genetics
  • Plasmids / metabolism
  • Protein Domains
  • Repressor Proteins / chemistry
  • Repressor Proteins / genetics
  • Repressor Proteins / metabolism*

Substances

  • Chromosomal Proteins, Non-Histone
  • Heterochromatin
  • Histones
  • Repressor Proteins
  • Chromobox Protein Homolog 5
  • Suv39h1 protein, mouse
  • Methyltransferases