1 School of Mathematics and Statistics and Evolution & Ecology Research Centre, University of New South Wales, Sydney, NSW, Australia. Electronic address: david.warton@unsw.edu.au.
2 Department of Mathematics and Statistics, McMaster University, Hamilton, Canada.
3 Biodiversity and Climate Research Centre, Frankfurt, Germany.
4 Metapopulation Research Center, Department of Biosciences, University of Helsinki, Finland; Centre for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology, Norway.
5 Department of Mathematics and Statistics, University of Jyväskylä, Jyväskylä, Finland.
6 Mathematical Sciences Institute, Australian National University, Canberra, Australia.