Stimulation of transforming growth factor-beta-1 and contact with type I collagen cooperatively facilitate irreversible transdifferentiation in proximal tubular cells

Biomed J. 2016 Feb;39(1):39-49. doi: 10.1016/j.bj.2015.08.004. Epub 2016 Mar 25.

Abstract

Background: By transdifferentiation, proximal tubular cells (PTC) have been considered as a source of interstitial myofibroblasts. We examined the combined effect of transforming growth factor-beta-1 (TGF-β1) stimulation and contact with type I collagen on PTC transdifferentiation.

Methods: Human kidney-2 cells were grown on type I substratum with the concurrent stimulation of TGF-β1.

Results: Following addition of TGF-β1, cells acquired an elongated fibroblastic appearance and an increase in α-smooth muscle actin (α-SMA) expression, a myofibroblastic marker. Upon addition of TGF-β1, E-cadherin expression, an epithelial marker, was reduced, while cytokeratin expression, another epithelial marker, remained unaltered. Following removal of TGF-β1, PTC regained an epithelial appearance and E-cadherin expression reverted to the unstimulated level, suggesting incomplete and reversible transdifferentiation. Addition of TGF-β1 to cells grown on type I collagen demonstrated a cooperatively increased α-SMA expression and decreased E-cadherin and cytokeratin expressions, suggesting more complete transdifferentiation. Co-stimulation of TGF-β1 and contact with type I collagen led to a stable cell phenotype and persistently decreased E-cadherin, which was not reversed upon removal of TGF-β1, indicating irreversible transdifferentiation. Addition of TGF-β1 or type I collagen caused a 4-fold increase in migratory cell number as compared to the control, whereas addition of both TGF-β1 and type I collagen led to an 11-fold increase.

Conclusions: TGF-β1 alone results in a reversible and incomplete transdifferentiation. The combination of TGF-β1 and exposure to type I collagen leads to an irreversible and complete PTC transdifferentiation.

Keywords: E-cadherin; Transdifferentiation; Transforming growth factor-beta-1; Type I collagen; α-Smooth muscle actin.

MeSH terms

  • Cadherins / metabolism
  • Cell Differentiation / drug effects*
  • Cell Differentiation / physiology
  • Cell Line
  • Cell Transdifferentiation / drug effects*
  • Cell Transdifferentiation / physiology
  • Collagen Type I / metabolism
  • Collagen Type I / pharmacology*
  • Epithelial Cells / cytology
  • Epithelial Cells / drug effects*
  • Fibroblasts / cytology
  • Fibroblasts / drug effects*
  • Humans
  • Transforming Growth Factor beta1 / metabolism
  • Transforming Growth Factor beta1 / pharmacology*

Substances

  • Cadherins
  • Collagen Type I
  • Transforming Growth Factor beta1