1 Department of Biology (M.H.v.C.-H., E.d.G., C.C.H.), div Cell Biology, Utrecht University, Utrecht, the Netherlands; Department of Neurology (M.H.v.C.-H, M.J.T., E.H., P.A.S.S.), Erasmus Medical Center, Rotterdam, the Netherlands; and Service of Neurology Hospital Clinic (L.S.), Institut d'Investigació Biomèdica August Pi i Sunyer (IDIBAPS), Barcelona, Spain.