1 Translational Gastroenterology Unit, Nuffield Department of Medicine, Experimental Medicine Division, John Radcliffe Hospital, University of Oxford , Oxford , UK.
2 MRC Human Immunology Unit, John Radcliffe Hospital, Weatherall Institute of Molecular Medicine, University of Oxford , Oxford , UK.
3 Translational Gastroenterology Unit, Nuffield Department of Medicine, Experimental Medicine Division, John Radcliffe Hospital, University of Oxford , Oxford , UK ; MRC Human Immunology Unit, John Radcliffe Hospital, Weatherall Institute of Molecular Medicine, University of Oxford , Oxford , UK.