1 Institute of Botany and Center for Molecular Bioscience Innsbruck, University of Innsbruck Innsbruck, Austria ; Department of Plant Biology and Ecology, University of the Basque Country (UPV/EHU) Bilbao, Spain.
2 Department of Plant Biology and Ecology, University of the Basque Country (UPV/EHU) Bilbao, Spain.
3 Institute of Agrobiotechnology, IdAB-CSIC-UPNA-Government of Navarra Pamplona, Spain.