The type I interferon response bridles rabies virus infection and reduces pathogenicity

J Neurovirol. 2011 Aug;17(4):353-67. doi: 10.1007/s13365-011-0041-6. Epub 2011 Jul 30.

Abstract

Rabies virus (RABV) is a neurotropic virus transmitted by the bite of an infected animal that triggers a fatal encephalomyelitis. During its migration in the nervous system (NS), RABV triggers an innate immune response, including a type I IFN response well known to limit viral infections. We showed that although the neuroinvasive RABV strain CVS-NIV dampens type I IFN signaling by inhibiting IRF3 phosphorylation and STAT2 translocation, an early and transient type I IFN response is still triggered in the infected neuronal cells and NS. This urged us to investigate the role of type I IFN on RABV infection. We showed that primary mouse neurons (DRGs) of type I IFN(α/β) receptor deficient mice (IFNAR(-/-) mice) were more susceptible to RABV than DRGs of WT mice. In addition, exogenous type I IFN is partially efficient in preventing and slowing down infection in human neuroblastoma cells. Intra-muscular inoculation of type I IFNAR deficient mice [IFNAR(-/-) mice and NesCre ((+/-)) IFNAR ((flox/flox)) mice lacking IFNAR in neural cells of neuroectodermal origin only] with RABV reveals that the type I IFN response limits RABV dissemination in the inoculated muscle, slows down invasion of the spinal cord, and delays mortality. Thus, the type I IFN which is still produced in the NS during RABV infection is efficient enough to reduce neuroinvasiveness and pathogenicity and partially protect the host from fatal infection.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Cell Line, Tumor
  • Female
  • Flow Cytometry
  • Humans
  • Immunohistochemistry
  • Injections, Intramuscular
  • Interferon Regulatory Factor-3 / genetics
  • Interferon Regulatory Factor-3 / metabolism
  • Interferon Type I* / biosynthesis
  • Interferon Type I* / immunology
  • Interferon Type I* / pharmacology
  • Mice
  • Mice, Knockout
  • Neuroblastoma / immunology
  • Neuroblastoma / pathology
  • Neuroblastoma / virology
  • Neurons / immunology*
  • Neurons / virology
  • Primary Cell Culture
  • Rabies / immunology*
  • Rabies / mortality
  • Rabies / pathology
  • Rabies / virology
  • Rabies virus / physiology*
  • Real-Time Polymerase Chain Reaction
  • Receptor, Interferon alpha-beta / deficiency*
  • Receptor, Interferon alpha-beta / genetics
  • Receptor, Interferon alpha-beta / immunology
  • STAT2 Transcription Factor / genetics
  • STAT2 Transcription Factor / metabolism
  • Signal Transduction / immunology*
  • Spinal Cord / immunology*
  • Spinal Cord / virology
  • Survival Rate
  • Viral Load / immunology

Substances

  • Interferon Regulatory Factor-3
  • Interferon Type I
  • Irf3 protein, mouse
  • STAT2 Transcription Factor
  • Stat2 protein, mouse
  • Receptor, Interferon alpha-beta