Abstract
Toll-like receptor-4 activation by lipopolysaccharide (LPS) induces the expression of interferon-beta (IFN-beta) in a MyD88-independent manner. Here we report that mice devoid of the JAK protein tyrosine kinase family member, Tyk2, were resistant to shock induced by high doses of LPS. Basal and LPS-induced expression of IFN-beta and IFN-alpha4 mRNA in Tyk2-null macrophages were diminished. However, Tyk2-null mice showed normal systemic production of nitric oxide and proinflammatory cytokines and the in vivo response to tumor necrosis factor (TNF) was unperturbed. IFN-beta-null but not STAT1-null mice were also resistant to high dose LPS treatment. Together, these data suggest that Tyk2 and IFN-beta are essential effectors in LPS induced lethality.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Cytokines / biosynthesis
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DNA-Binding Proteins / deficiency
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DNA-Binding Proteins / genetics
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DNA-Binding Proteins / metabolism
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Female
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Gene Expression
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Interferon Regulatory Factor-1
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Interferon Regulatory Factor-7
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Interferon Type I / genetics*
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Interferon-alpha / genetics
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Interferon-beta / genetics
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Lipopolysaccharides / toxicity
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Macrophage Activation
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Male
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Mice
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Mice, Inbred C57BL
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Mice, Knockout
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Nitric Oxide / biosynthesis
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Phosphoproteins / genetics
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Protein-Tyrosine Kinases / deficiency
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Protein-Tyrosine Kinases / genetics
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Protein-Tyrosine Kinases / metabolism*
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Proteins / genetics
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Proteins / metabolism*
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RNA, Messenger / genetics
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RNA, Messenger / metabolism
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STAT1 Transcription Factor
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Shock, Septic / etiology
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Shock, Septic / genetics
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Shock, Septic / immunology*
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TYK2 Kinase
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Trans-Activators / deficiency
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Trans-Activators / genetics
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Trans-Activators / metabolism
Substances
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Cytokines
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DNA-Binding Proteins
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Interferon Regulatory Factor-1
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Interferon Regulatory Factor-7
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Interferon Type I
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Interferon-alpha
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Irf1 protein, mouse
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Irf7 protein, mouse
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Lipopolysaccharides
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Phosphoproteins
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Proteins
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RNA, Messenger
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STAT1 Transcription Factor
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Stat1 protein, mouse
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Trans-Activators
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Nitric Oxide
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Interferon-beta
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Protein-Tyrosine Kinases
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TYK2 Kinase
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Tyk2 protein, mouse